NYMC Faculty Publications

LITAF, a BCL6 Target Gene, Regulates Autophagy in Mature B-cell Lymphomas

Author Type(s)

Faculty

Additional Author Affiliation

Touro College of Dental Medicine at NYMC

DOI

10.1111/bjh.12440

Journal Title

British Journal of Haematology

First Page

621

Last Page

630

Document Type

Article

Publication Date

9-1-2013

Department

Pathology, Microbiology and Immunology

Keywords

Autophagy, B-Lymphocyte Subsets, Cell Line, Tumor, DNA-Binding Proteins, Gene Expression Regulation, Neoplastic, Humans, Introns, Lymphoma, B-Cell, Neoplasm Proteins, Nuclear Proteins, Proto-Oncogene Proteins c-bcl-6, Transcription Factors, Transcription, Genetic, Tumor Necrosis Factor-alpha

Disciplines

Medicine and Health Sciences

Abstract

We have previously reported that LITAF is silenced by promoter hypermethylation in germinal centre-derived B-cell lymphomas, but beyond these data the regulation and function of lipopolysaccharide-induced tumour necrosis factor (TNF) factor (LITAF) in B cells are unknown. Gene expression and immunohistochemical studies revealed that LITAF and BCL6 show opposite expression in tonsil B-cell subpopulations and B-cell lymphomas, suggesting that BCL6 may regulate LITAF expression. Accordingly, BCL6 silencing increased LITAF expression, and chromatin immunoprecipitation and luciferase reporter assays demonstrated a direct transcriptional repression of LITAF by BCL6. Gain- and loss-of-function experiments in different B-cell lymphoma cell lines revealed that, in contrast to its function in monocytes, LITAF does not induce lipopolysaccharide-mediated TNF secretion in B cells. However, gene expression microarrays defined a LITAF-related transcriptional signature containing genes regulating autophagy, including MAP1LC3B (LC3B). In addition, immunofluorescence analysis co-localized LITAF with autophagosomes, further suggesting a possible role in autophagy modulation. Accordingly, ectopic LITAF expression in B-cell lymphoma cells enhanced autophagy responses to starvation, which were impaired upon LITAF silencing. Our results indicate that the BCL6-mediated transcriptional repression of LITAF may inhibit autophagy in B cells during the germinal centre reaction, and suggest that the constitutive repression of autophagy responses in BCL6-driven lymphomas may contribute to lymphomagenesis.

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